Robert W. Holley, Ph.D.

The Nobel Prize in Physiology or Medicine 1968
Nobel co-recipients: H. Gobind Khorana, Marshall W. Nirenberg

Organic Chemist. 'Interpreted genetic code and its function in protein synthesis' First chemical synthesis of penicillin. Research: amino acids, peptides, protein biosynthesis. Discovered and isolated tRNA [transfer RNA] - alanine transfer RNA.

Images

A team of Agriculture Research Service and Cornell University researchers, some of whom are pictured here, was the first to determine the molecular structure of RNA. This achievement won team leader Robert W. Holley (left), of ARS, the Nobel Prize in 1968. Author: Sol Goldberg. Photo: ARS Source: Wikimedia Commons.

Portrait of Robert William Holley working in the laboratory. Author: Smithsonian Institution. Source: Flickr: Portrait of Robert William Holley (1922-1993), Biochemist.

Robert W. Holley working. Source: ars.usda.gov

Robert W. Holley with his teen-age son. Source: ars.usda.gov

Photo of Robert W. Holley cropped from original. Author: Sol Goldberg. Photo Dept. of agriculture. Source: Wikimedia Commons

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