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Robert W. Holley, Ph.D.

The Nobel Prize in Physiology or Medicine 1968
Nobel co-recipients: H. Gobind Khorana, Marshall W. Nirenberg

Organic Chemist. 'Interpreted genetic code and its function in protein synthesis' First chemical synthesis of penicillin. Research: amino acids, peptides, protein biosynthesis. Discovered and isolated tRNA [transfer RNA] - alanine transfer RNA.

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A team of Agriculture Research Service and Cornell University researchers, some of whom are pictured here, was the first to determine the molecular structure of RNA. This achievement won team leader Robert W. Holley (left), of ARS, the Nobel Prize in 1968. Author: Sol Goldberg. Photo: ARS Source: Wikimedia Commons.

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Portrait of Robert William Holley working in the laboratory. Author: Smithsonian Institution. Source: Flickr: Portrait of Robert William Holley (1922-1993), Biochemist.

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Robert W. Holley working. Source: ars.usda.gov

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Robert W. Holley with his teen-age son. Source: ars.usda.gov

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Photo of Robert W. Holley cropped from original. Author: Sol Goldberg. Photo Dept. of agriculture. Source: Wikimedia Commons

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Holley
Photo Wiki. Painting Tim Tompkins - PaintHistory.com

Name: Robert William Holley
Birth: 28 January, 1922, Urbana, Il, USA
Death: 11 February, 1993, Los Gatos, CA, USA
Institution: Cornell University, Ithica, NY, USA
Award: "for their interpretation of the genetic code and its function in protein synthesis."
Portion of Cash: 1/3
Education: Public schools in Illinois, California, Idaho. Urbana High School 1938.
B. A. Chemistry, University of Illinois 1942.
Ph.D. organic chemistry, Cornell University, 1947. Mentor Professor Alfred T. Blomquist.
WWII interrupted graduate work 1944-1946; worked with Vincent du Vigneaud, Cornell University Medical College on first chemical synthesis of penicillin.
Postdoctoral Fellow, American Chemical Society, Mentor Professor Carl M. Stevens, Washington State University, 1947-1948.
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