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Robert W. Holley, Ph.D.

The Nobel Prize in Physiology or Medicine 1968
Nobel co-recipients: H. Gobind Khorana, Marshall W. Nirenberg

Organic Chemist. 'Interpreted genetic code and its function in protein synthesis' First chemical synthesis of penicillin. Research: amino acids, peptides, protein biosynthesis. Discovered and isolated tRNA [transfer RNA] - alanine transfer RNA.


His married Ann Dworkin in 1945. She taught Mathematics. Their son is Frederick.

Photo Wiki. Painting Tim Tompkins - PaintHistory.com

Name: Robert William Holley
Birth: 28 January, 1922, Urbana, Il, USA
Death: 11 February, 1993, Los Gatos, CA, USA
Institution: Cornell University, Ithica, NY, USA
Award: "for their interpretation of the genetic code and its function in protein synthesis."
Portion of Cash: 1/3
Education: Public schools in Illinois, California, Idaho. Urbana High School 1938.
B. A. Chemistry, University of Illinois 1942.
Ph.D. organic chemistry, Cornell University, 1947. Mentor Professor Alfred T. Blomquist.
WWII interrupted graduate work 1944-1946; worked with Vincent du Vigneaud, Cornell University Medical College on first chemical synthesis of penicillin.
Postdoctoral Fellow, American Chemical Society, Mentor Professor Carl M. Stevens, Washington State University, 1947-1948.
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Medal Cash and Philanthropy